2021 MNF TESP List.pdf

Malheur TES Plant Taxa by Habitat Type

Subalpine Wetland and Riparian Areas Scientific name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Anastrophyllum minutum

ANMI8 tiny notchwort liverwort

Peaty soil, at high elevations. Associated with other bryophytes in tight mats. On wet soil, rocky ledges, or on wet rocks in sub alpine or alpine zones. Usually on north-facing exposures.

Barbilophozia lycopodioides

BALY giant fourpoint, maple liverwort

Peaty soil or decomposed organic detritus on damp ledges of rock outcrops and cliffs at higher elevations. Acid soil or soil-covered rocks. Not necessarily on mineral soil.

Sites receive abundant snowfall.

Harpanthus flotovianus

HAFL9 great mountain flapwort liverwort

Bogs and fens; mostly on humus, soil covered rocks, and decaying wood in forests.

Site on Umatilla NF is in a sub-alpine meadow, surrounded by subalpine fir and spruce forest.

Jungermannia polaris

JUPO3 Arctic flapwort

(liverwort)

In permanent seeps, thin soil over calcareous rocks. In ultraoligotrophic lakes. Forming mats or hidden among other bryophytes, over peaty soils on damp ledges and crevices of rocks, sometimes along streams, or aquatic. Partial shade to full exposure.

Peltolepis quadrata

PEQU7

shieldscale liverwort

Forming small mats on moist, exposed to shady soil or on ledges and crevices on talus slopes and outcrops. Usually calciphile, also igneous & ultramafic rocks. Cold forest to white bark pine types.

Preissia quadrata

PRQU2

blister ribbon, narrow mushroom-headed liverwort

On soil w/little organic matter, typically on ledges or cliffs. On MAL NF in peaty fen.

Shaded creeks, seeps, splash zones. Terrestrial, on damp mineral soil; reportedly a strict calciphile, but not in Columbia Gorge. In thin soil over calcareous rocks.

Pseudocalliergon trifarium

PSTR5 blunt water moss, worm moss

Forming lawns or intermixed with other bryophytes in medium to rich montane fens;

grows submerged to emergent in pools or on saturated ground; usually in full sunlight.

Fen pools may dry up in late summer.

Ptilidium pulcherrimum

PTPU2 blunt water moss, Forming low, dense mats at the base of trees, on decaying wood, among boulders in talus, ledges of cliffs, and rarely on soil, in cool moist habitats. Perennially moist tree bases, rotten logs, and on rocks, cliffs, and ledges.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Tortula mucronifolia

TOMU70 mucron-leaf tortula moss

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Vasculars

Botrychium ascendens crenulatum hesperium lineare paradoxum pedunculosum

BOAS2

BOCR

BOHE5

BOLI7

BOPA9

BOPE4

moonworts

Moist meadows, riparian zones, moist roadsides, openings in cold forests. Often in calcareous soils, but not always. Lower montane, mesic coniferous forest, and grassy fields. Often on the drier edges of wet meadows.

Botrychium lunaria BOLU common moonwort Open fields, occasionally forest in southern occurrences. Moist meadows, riparian zones, moist roadsides, openings and edges of cold forests.

Botrychium montanum

BOMO

mountain moonwort

Dark, coniferous forests, usually near swamps and streams. Wet meadows, saturated soils. Often growing in a bed of mosses. This species tends to grow in wetter sites than the other Botrychiums.

Carex atrosquama

CAAT8

brass-fruit sedge

Alpine and supalpine grasslands, moist to mesic meadows, lake shorelines, and streamsides. Documented populations in Wallowas are mostly at base of limestone cliffs.

Carex lasiocarpa var. americana CALA11 slender woolly sedge Fens, bogs, lakeshores, streambanks, seepage areas, and wet meadows. In deep, organic acid soils with low to moderate nutrients. Sometimes forming floating mats.

Carex media CAME9 Scandinavian sedge

Montane to subalpine meadows, streambanks, seepage areas, bog margins. Mossy, moist forests.

Carex micropoda

CAMI16

timberline sedge

Moist alpine meadows and snowmelt basins, areas irrigated by meltwater. In cracks between rocks on talus fields near alpine lakes. Mainly where heavy snow cover protects from extreme cold temperatures, prevents competition, and provides summer moisture.

Carex pelocarpa CAPE5 dusky-seed sedge Dry alpine slopes, ridges, boulder fields, talus slopes, and on rocky stream sides in damp but well-drained soils, usually with no competing vegetation.

Carex saxatilis CASA10 limestone or russet sedge

Fens, bogs, lakeshores, wet tundra, roadside ditches, ponds, and slow moving streams; often in shallow water. Always at high elevations. In areas with peaty soils.

Carex scirpoidea ssp. stenochlaena CASCS9 Alaskan single-spiked sedge Moist meadows, streambanks, and rocky slopes. Beside waterfalls and on seepy cliffs.

Somewhat acidic substrates.

subnigricans CASU7 dark mountain sedge Moist rocky slopes and meadows, subalpine to alpine areas.

Carex vernacula CAVE5 foetid or native sedge

Alpine and subalpine wet meadows, rocky slopes, edges of headwater streams and lakeshores. Moist forest openings just below treeline.

Comastoma tenellum

COTE13

Slender gentian

Subalpine to alpine meadows. Never found directly on hummocks. Associated species in Washington include Salix planifolia, Vaccinium caespitosum, Carex spp., Dasiphora fruticosa, and mosses. At Steens mtn w/Carex aurea and Botrychiums.

Cryptogramma stelleri

CRST2

Steller's rock-brake

Sheltered calcareous cliff crevices and rock ledges, typically in coniferous or boreal forest. On wet cliffs, and ledges, along streams, under waterfalls. Limestone substrate.

Shady sites. Usually north-facing aspects.

Diphasiastrum complanatum LYCO3 ground-cedar Dry open coniferous or mixed dry to wet forest slopes, or in riparian areas. Often on rotting logs, or in thick duff. Reportedly also in meadows and on open ridge tops.

Kobresia myosuroides KOMY Bellard's kobresia Tundra, grassland, heaths, bare rocky, dry to wet ground. Moist meadows, seeps, riparian areas. Often above timberline; high elevations only.

Lophozia gillmanii

LOGI3 Gillman's pawwort

On peaty soil, usually on cliffs or ledges; obligate calciphile? Mossy, constantly-irrigated basic rock ledges and in mossy base-rich flushes and rocks by streams. On granite derived substrates on the Umatilla NF.

Salix farriae SAFA Farr's willow Wet meadows, stream banks, moderate to high elevations.

Salix wolfii SAWOI2 Wolf's willow Riparian, wet meadows, waterfalls, stream-sides, moist shrub lands; moderate to high elevations.

Saxifraga adscendens ssp.

oregonensis

SAADO2 wedge-leaved saxifrage

Rock crevices, glacial moraines, alpine meadows, along streams. Damp cliffs where soil has accumulated. High elevations only. Sites in Wallowas are on limestone, reported also on quartzite and gneiss in Idaho. Needs some amount of moisture.

Swertia perennis SWPE Felwort Subalpine wet areas, stream sides, springs, wet meadows, wet alpine slopes Bogs.

Not clear if in peatlands. Moderate to high elevation in the mountains.

Trollius albiflorus TRLAA2 American globeflower

Open, wet places, more or less acidic soils, montane to alpine. Seeps, springs, and vernally wet swales in spruce/fir forest.

Subalpine Dry Areas name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Peaty soil, at high elevations. Associated with other bryophytes in tight mats. On wet soil, rocky ledges, or on wet rocks in sub alpine or alpine zones. Usually north-facing exposures.

Peaty soil or decomposed organic detritus on damp ledges of rock outcrops and cliffs at higher elevations. Acid soil or soil-covered rocks. Not necessarily on mineral soil.

Sites receive abundant snowfall.

Bryum calobryoides BRCA20 Bryum moss On protected to exposed rock at moderate to high elevations. Can also grow on moist soil.

Encalypta brevipes

ENBR2

candle snuffer moss, stubby extinguisher moss

Soil on ledges and in crevices on cliffs, igneous and siliceous substrates. Sites may be subject to frequent fog penetration.

Bogs and fens; mostly on humus, soil covered rocks, and decaying wood in forests.

Site on Umatilla NF is in a sub-alpine meadow, surrounded by subalpine fir and spruce forest.

JUPO3

Arctic flapwort

In permanent seeps, thin soil over calcareous rocks. In ultraoligotrophic lakes.

Forming mats or hidden among other bryophytes, over peaty soils on damp ledges and crevices of rocks, sometimes along streams, or aquatic. Partial shade to full exposure.

On peaty soil, usually on cliffs or ledges; obligate calciphile? Mossy, constantly-irrigated basic rock ledges and in mossy base-rich flushes and rocks by streams. On granite derived substrates on the Umatilla NF.

Peltolepis quadrata

PEQU7

shieldscale liverwort

Forming small mats on moist, exposed to shady soil or on ledges and crevices on talus slopes and outcrops. Usually calciphile, also igneous & ultramafic rocks. Cold forest to white bark pine types.

narrow mushroom-headed liverwort

On soil w/little organic matter, typically on ledges or cliffs. On MAL NF in peaty fen.

Shaded creeks, seeps, splash zones. Terrestrial, on damp mineral soil; reportedly a strict calciphile, but not in Columbia Gorge. In thin soil over calcareous rocks.

Forming low, dense mats at the base of trees, on decaying wood, among boulders in talus, ledges of cliffs, and rarely on soil, in cool moist habitats. Perennially moist tree bases, rotten logs, and on rocks, cliffs, and ledges.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Vasculars

Boechera paddoensis BOPA16 Mt. Adams rockcress Alpine and subalpine rocky ridges, 1500-2100 m.

Botrychium ascendens crenulatum hesperium lineare paradoxum pedunculosum

BOAS2

BOCR

BOHE5

BOLI7

fields. Often on the drier edges of wet meadows.

Botrychium lunaria BOLU common moonwort Open fields, occasionally forest in southern occurrences. Moist meadows, riparian zones, moist roadsides, openings and edges of cold forests.

Carex media CAME9 Scandinavian sedge

Montane to subalpine meadows, streambanks, seepage areas, bog margins. Mossy, moist forests.

Carex micropoda

CAMI16

timberline sedge

Moist alpine meadows and snowmelt basins, areas irrigated by meltwater. In cracks between rocks on talus fields near alpine lakes. Mainly where heavy snow cover protects from extreme cold temperatures, prevents competition, and provides summer moisture.

Carex nardina CANA2 spikenard sedge Dry alpine ridges and slopes, abrasion plateaus, dry tundra, heaths, and fellfields.

Usually at, or above, timberline.

Carex pelocarpa CAPE5 dusky-seed sedge Dry alpine slopes, ridges, boulder fields, talus slopes, and on rocky streamsides in damp but well-drained soils, usually w/no competing vegetation.

Carex subnigricans CASU7 dark mountain sedge Moist rocky slopes and meadows, subalpine to alpine areas.

Carex tahoensis CATA Tahoe sedge Dry subalpine and alpine ridges and grasslands.

Carex vernacula CAVE5 foetid or native sedge

Alpine and subalpine wet meadows, rocky slopes, edges of headwater streams and lakeshores. Moist forest openings just below treeline.

Castilleja viscidula

CAVI9 Sticky Indian paintbrush

Mid-elevation juniper and mixed conifer woodlands to rocky subalpine slopes and summits. Associated species: Pinus albicaulis, Ipomopsis aggregata, Eriogonum umbellatum, E. sphaerocephalum and Eremogone (Arenaria) spp.

Cheilanthes feei CHFE Fee's lip fern Cliff crevices, especially on limestone. On dry limestone influenced cliffs (sometimes basalt), from low elevations in deep canyons to high in sub-alpine.

Cymopterus nivalis

CYNI3 snowline cymopterus

Open rocky places. Moderate to high elevations. Found on volcanic tuffs and bald areas with less than 30% vegetation cover. Artemisia arbuscula is a common associate.

Diphasiastrum complanatum LYCO3 ground-cedar Dry open coniferous or mixed dry to wet forest slopes, or in riparian areas. Often on rotting logs, or in thick duff. Reportedly also in meadows and on open ridge tops.

Kobresia myosuroides KOMY Bellard's kobresia Tundra, grassland, heaths, bare rocky, dry to wet ground. Moist meadows, seeps, riparian areas. Often above timberline; high elevations only.

Lomatium tarantuloides

LOTA4

spider biscuit-root

Clearings in upper montane subalpine fir-dominated forest; Primarily in shallow, gravelly, serpentine-derived lithosolic soils on flat to gently sloping southerly to easterly aspects; sites are ephemerally moist to wet and generally poorly vegetated, with L. tarantuloides tending to be the dominant vascular plant species.

Phlox hendersonii PHHE9 Henderson's phlox Alpine fell fields, open gravely slopes and ridges. Glacial moraines. Exposed, rocky places.

Pinus albicaulis PIAL whitebark pine Subalpine, usually near timberline. Usually fairly dry sites with thin, rocky, cold soils.

Piptatheropsis exigua PIEX4 Little mountain ricegrass Rocky slopes and outcrops in upper montane regions. 1200– 2800 m

Potentilla versicolor var.

darachii

!!POVED

Darrach’s cinquefoil

Ultramafic substrates in subalpine to alpine zones, particularly on ‘sky islands’. Often near outcrops and/or with abundant cobble-size rocks which are moist early in the season but eventually dry out.

Rhodiola integrifolia ssp.

integrifolia

RHINI

Ledge stonecrop

Rocky slopes, cliffs, alpine meadows.

Salix nivalis SANI8 Snow willow Alpine tundra, cirques, lake basins, rocky slopes and ridges, fellfields.

Saxifraga adscendens ssp.

oregonensis

SAADO2 wedge-leaved saxifrage

Rock crevices, glacial moraines, alpine meadows, along streams. Damp cliffs where soil has accumulated. High elevations only. Sites in Wallowas are on limestone, reported also on quartzite and gneiss in Idaho. Needs some amount of moisture.

Cold and Moist Upland Coniferous Forests name

Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Albatrellus avellaneus ALAV Avel’s albetrellus polyphore Fruits infrequently in the fall on soil in moist coniferous forest. Associated with hemlock, Douglas-fir and spruce.

Peaty soil or decomposed organic detritus on damp ledges of rock outcrops and cliffs at higher elevations. Acid soil or soil-covered rocks. Not necessarily on mineral soil.

Sites receive abundant snowfall.

Bryum calobryoides BRCA20 Bryum moss On protected to exposed rock in mid to upper elevation where it can also grow on moist soil.

Forming low, dense mats at the base of trees, on decaying wood, among boulders in talus, ledges of cliffs, and rarely on soil, in cool moist habitats. Perennially moist tree bases, rotten logs, and on rocks, cliffs, and ledges.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

TOMO70 mucron-leaf tortula

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Tritomeria exsecta TREX4 Cut notchwort Damp, mossy rocks and rotting logs with other bryophytes in spruce-fir forests.

Vasculars

Botrychium ascendens crenulatum hesperium lineare paradoxum pedunculosum

BOAS2

BOCR

BOHE5

BOLI7

fields. Often on the drier edges of wet meadows.

Botrychium lunaria BOLU common moonwort Open fields, occasionally forest in southern occurrences. Moist meadows, riparian zones, moist roadsides, openings and edges of cold forests.

soils. Often growing in a bed of mosses. This species tends to grow in wetter sites than the other Botrychiums.

Cypripedium fasciculatum CYFA clustered lady's-slipper Grand fir to ponderosa pine, and warm riparian forests; w/sub-alpine fir in Colorado.

Generally in 60-100% shade. Often w/mountain lady's slipper. In damp to dry sites.

Diphasiastrum complanatum LYCO3 ground-cedar Dry open coniferous or mixed dry to wet forest slopes, or in riparian areas. Often on rotting logs, or in thick duff. Reportedly also in meadows and on open ridge tops.

Listera borealis

LIBO4

northern twayblade

In moist, rich humus of mossy coniferous forest, swamps, often along cold streams, acidic soils. Most known sites are in older forests. Associated tree species include spruce, true firs, and Doug fir. Moderate elevations.

Trollius albiflorus TRLAA2 American globeflower

Dry Upland Coniferous Forests and Upland Woodlands

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Texosporium sancti-jacobi

TESA

woven spore lichen

Arid to semi-arid shrub-steppe, grassland or savannah. Warm, dry sites w/silty or rocky soils. On flat to gentle north facing slopes in arid areas. Often on decomposing bunchgrass bases, dead Selaginella, or on mammal scat (rabbit).

Vasculars

Achnatherum nevadense ACNE10 Nevada needlegrass Montane to subalpine sagebrush steppe and open woodlands.

Astragalus diaphanus var.

diurnus

ASDID4

John Day milkvetch

Shallow gravelly soil over basalt, sandbars, or sandy banks of intermittent rivers, often in juniper woodlands. Associated species: Artemisia tridentata, A. rigida, Eriogonum compositum, Eriogonum phaerocephalum, Eriophyllum lanatum, Juniperus occidentalis, Ericameria nauseosa, Lewisia rediviva, and Salvia dorii.

Astragalus misellus var.

misellus

ASMIM3

pauper milkvetch

Sagebrush and western juniper slopes on sandy and loamy, often rocky soils.

Associated species may include: juniper, big, low, and stiff sagebrushes, Thurber’s needlegrass, and Sandberg’s bluegrass.

Astragalus tegetarioides

ASTE4 Deschutes milkvetch

Dry pine forest, sagebrush flats, on volcanic ash or basaltic substrates, cracks of welded tuff outcrops. In transition zone between sagebrush and ponderosa pine forest. Endemic to Emigrant Creek Ranger District (and west).

Botrychium campestre BOCA5 Iowa moonwort Extremely inconspicuous in prairies, dunes, grassy railroad sidings, and fields over limestone.

CalyptridIum roseum

CARO

rosy calyptridium

Sagebrush desert to montane coniferous forest. Gravely to sandy soils, often in low moist swales among sagebrush. Often in alkaline soils. In areas w/low cryptogram cover. Historic Malheur NF site is in dry ponderosa pine site (wetter microsite?).

Carex cordillerana CACO81 cordilleran sedge Riparian terraces, mesic forests, with grand fir and Douglas fir. Aspen forests. Rocky slopes, in leaf litter and duff. Also in rocky areas with juniper and mountain mahogany.

Castilleja viscidula

CAVI9 Sticky Indian paintbrush

Mid-elevation juniper and mixed conifer woodlands to rocky subalpine slopes and summits. Associated species: Pinus albicaulis, Ipomopsis aggregata, Eriogonum umbellatum, E. sphaerocephalum and Eremogone (Arenaria) spp.

Cryptantha simulans CRSI2 Pine woods cryptantha In open ponderosa pine forests. Reported on open rocky slope at Baldy Mtn, in Elkhorns, on Wallowa-Whitman NF.

Eriogonum salicornioides

ERSA8 playa or saltwort buckwheat

On sparsely vegetated, open, ash deposits and clayey substrates, sometimes in disturbed areas. Saltbush, greasewood, and sagebrush communities; juniper woodlands possibly surrounding. Only suspected on Emigrant Creek Ranger District.

Ipomopsis tenuituba ssp.

tenuituba

IPTET2 Slendertube skyrocket

Open, rocky, dry slopes. In Blue Mountains at relatively high elevations, >6,000 ft.

Luina serpentina LUSE colonial luina Open, rocky sites with poor soil development. Usually on steep slopes, above small tributaries. In dry shrub, open juniper, ponderosa pine, and Douglas fir forest areas.

Muhlenbergia minutissima

MUMI2 annual or little-seed muhly grass

Sandy soils on riverbanks, moist meadows, pond shores and edges of reservoirs.

Open, rocky and apparently dry slopes also. Open, more or less disturbed, sandy slopes, and seeps.

Penstemon deustus var.

variabilis

PEDEV2 Variable hot rock penstemon

Dry foothills and lowlands, on open, dry, thin soils over basalt. Associated species include Pseudoroegneria spicata, Festuca idahoensis, Eriogonum compositum, Eriophyllum lanatum, Astragalus spp., Madia gracilis, and Collomia grandiflora.

Phacelia

PHMI7 least or dwarf phacelia

Moist, open places, streambanks, meadows, ephemerally moist swales. Vernally moist openings in ponderosa pine or Doug fir forest. Often with Veratrum. In aspen in

WA.

Phlox multiflora

PHMU3 many-flowered phlox

Basalt cliffs, rocky outcrops and bluffs, rocky openings and wooded rocky areas in dry forest. Sometimes on loose substrate as well as cracks in cliffs and rock outcrops.

Highest elevation in OR is 4,800 ft., however found higher up in Rocky Mountains.

Thelypodium eucosmum

THEU arrow-leaf thelypody

Under and around western juniper, ponderosa pine, and Doug fir trees; in canyons, seasonal creek drainages, and seeps/springs. Also found in vernally moist areas in ponderosa pine forests and in sage. Restricted to serpentine and ultramafic soils.

Lithosols (Scablands) rocky soils. On flat to gentle north facing slopes in arid areas. Often on decomposing bunchgrass bases, dead Selaginella, or on mammal scat (rabbit).

Vasculars

Achnatherum hendersonii ACHE10 Henderson's ricegrass Basalt lithosols; shallow rocky soils, sometimes w/stiff sage, strict buckwheat, and Ponderosa pine surrounding. Can be in deeper-soiled areas than Wallowa ricegrass.

Achnatherum wallowaense ACWA4 Wallowa ricegrass Basalt lithosols; shallow rocky soils, sometimes w/stiff sage, strict buckwheat, and

Ponderosa pine surrounding.

Astragalus in juniper woodlands. Associated species: Artemisia tridentata, A. rigida, Eriogonum compositum, Eriogonum phaerocephalum, Eriophyllum lanatum, Juniperus occidentalis, Ericameria nauseosa, Lewisia rediviva, and Salvia dorii.

Eleocharis bolanderi

ELBO

Bolander's spike-rush

Vernally wet swales, along intermittent streams, wet depressions in moist meadows and lithosols. In slight depressions that hold snow later in the season than surrounding areas. Surrounding forest is usually ponderosa pine. Habitat and plants are dry by mid-summer.

Eremothera pygmaea CAPY5 dwarf evening-primrose Dry plains and slopes with unstable soils or in gravel in steep talus, dry washes, banks, and road cuts. Open, bare ground. Sometimes w/big sage or bitterbrush.

cusickii

ERCU3

Cusick's buckwheat

Rocky sagebrush desert. Barren flats and hills w/dry soil over weathered basalt and welded tuff. Associated species include low sage and juniper. Only suspected on Emigrant Creek RD.

Barren Unusual Soils name Code Common name Habitat details

Vasculars

Chaenactis xantiana CHXA desert chaenactis Dry, open places, deep, loose sandy soil or on dunes, less often on gravelly soil. In sagebrush steppe areas. Only suspected for Emigrant Creek RD.

Eriogonum cusickii ERCU3 Cusick's buckwheat Rocky sagebrush desert. Barren flats and hills w/dry soil over weathered basalt and welded tuff. Associated species include low sage and western juniper.

On sparsely vegetated, open, ash deposits and clayey substrates, sometimes in disturbed areas. Saltbush, greasewood, and sagebrush communities; juniper woodlands possibly surrounding.

Heliotropium curassavicum HECU3 salt heliotrope Wet ground, saline flats, and alkaline soils. Dried edges and centers of vernal pools.

Low to moderate elevations. Probably only on Emigrant Creek RD.

Lupinus lepidus var. cusickii LUCUC2 Cusick's lupine Residual soils from ancient lakebeds, volcanic ash and clay soils. Barren tuffaceous or gravelly clay bluffs and gullied riverbanks. Plants often in eroded drainage channels.

Myosurus sessilis MYSE2 short-stemmed mousetail Alkali flats. Dry pool beds. Vernal pools within grasslands. Clay bottom vernal pools.

Phlox hendersonii PHHE9 Henderson’s phlox Alpine fell fields, open gravely slopes, ridges. Glacial moraines. High elevations.

Polygonum gabrielae !POGA Gabriel’s knotweed Dry, barren, gravelly, serpentine slopes.

Stanleya confertiflora

STCO2

biennial stanleya

Dry plains and low sand or clay hills. On barren clay and diatomaceous soils in sagebrush habitats. Tuffaceous sedimentary rocks on geology layer. Sometimes with juniper.

THEU

arrow-leaf

Under and around western juniper, ponderosa pine, and Doug fir trees; in canyons, seasonal creek drainages, and seeps/springs. Also found in vernally moist areas in ponderosa pine forests and in sage. Restricted to serpentine and ultramafic derived soils on Malheur NF.

Sagebrush Steppe rocky soils. On flat to gentle north facing slopes in arid areas. Often on decomposing bunchgrass bases, dead Selaginella, or on mammal scat (rabbit).

Vasculars

Achnatherum nevadense ACNE10 Nevada needlegrass Montane to subalpine sagebrush steppe and open woodlands.

Astragalus in juniper woodlands. Associated species: Artemisia tridentata, A. rigida, Eriogonum compositum, Eriogonum phaerocephalum, Eriophyllum lanatum, Juniperus occidentalis, Ericameria nauseosa, Lewisia rediviva, and Salvia dorii.

Astragalus misellus var.

misellus

ASMIM3

pauper milkvetch

Sagebrush and western juniper slopes on sandy and loamy, often rocky soils.

Associated species may include: juniper, big, low, and stiff sagebrushes, Thurber’s needlegrass, and Sandberg’s bluegrass.

Astragalus tegetarioides

ASTE4 Deschutes milkvetch

Dry pine forest, sagebrush flats, on volcanic ash or basaltic substrates, cracks of welded tuff outcrops. In transition zone between sagebrush and ponderosa pine forest. Endemic to Emigrant Creek Ranger District (and west).

Calyptridum roseum

CARO

rosy calyptridium

Sagebrush desert to montane coniferous forest. Gravely to sandy soils, often in low moist swales among sagebrush. Often in alkaline soils. In areas w/low cryptogram cover. Historic Malheur NF site is in dry ponderosa pine site (wetter microsite?).

Chaenactis xantiana CHXA desert chaenactis Dry, open places, deep, loose sandy soil or on dunes, less often on gravelly soil. In sagebrush steppe areas. Only suspected for Emigrant Creek RD

Vernally wet swales, along intermittent streams, wet depressions in moist meadows and lithosols. In slight depressions that hold snow later in the season than surrounding areas. Surrounding forest is usually ponderosa pine. Habitat and plants are dry by mid-summer.

Eremothera pygmaea CAPY5 dwarf evening-primrose Dry plains and slopes with unstable soils or in gravel in steep talus, dry washes, banks, and road cuts. Open, bare ground. Sometimes w/big sage or bitterbrush.

Eriogonum cusickii ERCU3 Cusick's buckwheat Rocky sagebrush desert. Barren flats and hills w/dry soil over weathered basalt and welded tuff. Associated species include low sage and western juniper.

On sparsely vegetated, open, ash deposits and clayey substrates, sometimes in disturbed areas. Saltbush, greasewood, and sagebrush communities; juniper woodlands possibly surrounding.

Stanleya confertiflora

STCO2

biennial stanleya

Dry plains and low sand or clay hills. On barren clay and diatomaceous soils in sagebrush habitats. Tuffaceous sedimentary rocks on geology layer. Sometimes with juniper.

Under and around western juniper, ponderosa pine, and Doug fir trees; in canyons, seasonal creek drainages, and seeps/springs. Also found in vernally moist areas in ponderosa pine forests and in sage. Restricted to serpentine and ultramafic rocks on Malheur NF.

Low to Moderate Elevation Cliffs, Rock Outcrops and Talus name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Bryum calobryoides BRCA20 Bryum moss On protected to exposed rock at moderate to high elevations. Can also grow on moist soil.

Encalypta brevipes

ENBR2

candle snuffer moss, stubby extinguisher moss

Soil on ledges and in crevices on cliffs, igneous and siliceous substrates. Sites may be subject to frequent fog penetration.

Entosthodon fascicularis

ENFA2

banded cord-moss

On seasonally wet, exposed soil in seeps, or along intermittent streams. Grasslands, grassy balds, and rock outcrops. Usually hidden among grasses, other mosses, and litter.

In permanent seeps, thin soil over calcareous rocks. In ultraoligotrophic lakes.

Forming mats or hidden among other bryophytes, over peaty soils on damp ledges

On peaty soil, usually on cliffs or ledges; obligate calciphile? Mossy, constantly-irrigated basic rock ledges and in mossy base-rich flushes and rocks by streams. On granite derived substrates on the Umatilla NF.

narrow mushroom-headed liverwort

On soil w/little organic matter, typically on ledges or cliffs. On MAL NF in peaty fen.

Shaded creeks, seeps, splash zones. Terrestrial, on damp mineral soil; reportedly a strict calciphile, but not in Columbia Gorge. In thin soil over calcareous rocks.

Forming low, dense mats at the base of trees, on decaying wood, among boulders in talus, ledges of cliffs, and rarely on soil, in cool moist habitats. Perennially moist tree bases, rotten logs, and on rocks, cliffs, and ledges.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Vasculars cordillerana

CACO81

cordilleran sedge

Riparian terraces, mesic forests, with grand fir and Douglas fir. Aspen forests. Rocky slopes, in leaf litter and duff. In rocky areas with juniper and mountain mahogany on Malheur NF. Near ephemeral streams in steep ravines in Hells Canyon NRA.

Carex scirpoidea ssp. stenochlaena CASCS9 Alaskan single-spiked sedge Moist meadows, streambanks, and rocky slopes. Beside waterfalls, and on seepy cliffs. Somewhat acidic substrates.

Cheilanthes feei CHFE Fee's lip fern Ciiff crevices, especially on limestone. On dry limestone influenced cliffs (sometimes basalt), from low elevations in deep canyons to high in sub-alpine.

Cryptogramma stelleri

CRST2

Steller's rock-brake

Sheltered calcareous cliff crevices and rock ledges, typically in coniferous or boreal forest. On wet cliffs, and ledges, along streams, under waterfalls. Limestone substrate. Shady sites. Usually north-facing aspects.

Eremothera pygmaea CAPY5 dwarf evening-primrose Dry plains and slopes with unstable soils or in gravel in steep talus, dry washes, banks, and road cuts. Open, bare ground. Sometimes w/big sage or bitterbrush.

Luina serpentina LUSE colonial luina Open, rocky sites with poor soil development. Usually on steep slopes, above small tributaries. In dry shrub, open juniper, ponderosa pine, and Douglas fir forest areas.

Sandy soils on riverbanks, moist meadows, pond shores and edges of reservoirs.

Open, rocky and apparently dry slopes also. Open, more or less disturbed, sandy slopes, and seeps.

Pellaea bridgesii PEBR5 Bridge's cliff-brake Rocky slopes and cliffs, on granitic substrate. Talus slopes, scree, rock outcrops;

often on southern exposures on argillite in Wallowa and Blue Mountains.

Phlox multiflora

PHMU3 many-flowered phlox

Basalt cliffs, rocky outcrops and bluffs, rocky openings and wooded rocky areas in dry forest. Sometimes on loose substrate as well as cracks in cliffs and rock outcrops.

Highest elevation in OR is 4,800 ft., however found higher up in Rocky Mountains.

Cold Riparian Forests, Shrublands, Springs and Wetlands (Excluding Peatland-Only Species)

Scientific name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Albatrellus avellaneus ALAV Avel’s albetrellus polyphore Fruits infrequently in the fall on soil in moist coniferous forest. Associated with hemlock, Douglas-fir and spruce.

Peaty soil, at high elevations. Associated with other bryophytes in tight mats. On wet soil, rocky ledges, or on wet rocks in sub alpine or alpine zones. Usually on north-facing exposures.

Entosthodon banded cord-moss, Hasselquist’s hyssop

On seasonally wet, exposed soil in seeps, or along intermittent streams. Grasslands, grassy balds, and rock outcrops. Usually hidden among grasses, other mosses, and litter.

Bogs and fens; mostly on humus, soil covered rocks, and decaying wood in forests.

Site on Umatilla NF is in a sub-alpine meadow, surrounded by subalpine fir and spruce forest.

In permanent seeps, thin soil over calcareous rocks. In ultraoligotrophic lakes.

Forming mats or hidden among other bryophytes, over peaty soils on damp ledges

On peaty soil, usually on cliffs or ledges; obligate calciphile? Mossy, constantly-irrigated basic rock ledges and in mossy base-rich flushes and rocks by streams. On granite derived substrates on the Umatilla NF.

Preissia quadrafeta narrow mushroom-headed liverwort

On soil w/little organic matter, typically on ledges or cliffs. On MAL NF in peaty fen.

Shaded creeks, seeps, splash zones. Terrestrial, on damp mineral soil; reportedly a strict calciphile, but not in Columbia Gorge. In thin soil over calcareous rocks.

Forming low, dense mats at the base of trees, on decaying wood, among boulders in talus, ledges of cliffs, and rarely on soil, in cool moist habitats. Perennially moist tree bases, rotten logs, and on rocks, cliffs, and ledges.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Vasculars

Botrychium ascendens crenulatum hesperium lineare paradoxum pedunculosum

BOAS2

BOCR

BOHE5

BOLI7

fields. Often on the drier edges of wet meadows.

BOMO mountain moonwort

Dark, coniferous forests, usually near swamps and streams. Wet meadows, saturated soils. Often growing in a bed of mosses. This species tends to grow in wetter sites than the other Botrychiums.

Carex retrorsa CARE4 retrorse sedge Floodplains in forests, streamsides, lakeshores, swamps, wet thickets, wet meadows.

Sandy riverbanks. Sites are wet in spring, but may dry out by late summer.

Carex scirpoidea ssp. stenochlaena CASCS9 Alaskan single-spiked sedge Moist meadows, streambanks, and rocky slopes. Beside waterfalls and on seepy cliffs. Somewhat acidic substrates.

Cypripedium fasciculatum

CYFA clustered lady's-slipper

Grand fir to ponderosa pine, and warm riparian forests; w/sub-alpine fir in CO.

Generally found in 60-100% shade. Often w/mtn. lady's slipper. In damp to dry sites.

Seeps/springs, riparian zones.

Diphasiastrum complanatum LYCO3 ground-cedar Dry open coniferous or mixed dry to wet forest slopes, or in riparian areas. Often on rotting logs, or in thick duff. Reportedly also in meadows and on open ridge tops.

Epilobium palustre EPPA marsh willowherb Wet meadows, shorelines, peatlands, streambanks, marshes, seeps.

Listera borealis

LIBO4

northern twayblade

In moist, rich humus of mossy coniferous forest, swamps, often along cold streams, acidic soils. Most known sites are in older forests. Associated tree species include spruce, true firs, and Doug fir. Moderate elevations.

Ophioglossum pusillum

OPPU3

adder's-tongue

Seasonally wet areas in pastures, old fields, roadside ditches, bogs, fens, wet meadows, flood plains, moist woods, grassy swales, dry or damp sand, dry hillsides, seasonally wet, acidic soil.

Salix farriae SAFA Farr’s willow Wet meadows, stream banks, moderate to high elevations.

Salix wolfii SAWOI2 Wolf's willow Riparian, wet meadows, waterfalls, stream-sides, moist shrub lands; moderate to high elevations.

Trifolium douglasii TRDO Douglas' clover Moist or mesic meadows, prairie remnants, along riparian areas along streams. In swales, along intermittent streams, and in vernally wet areas.

Trollius albiflorus TRLAA2 American globeflower

Aquatic Species name Code Common name Habitat details

In permanent seeps, thin soil over calcareous rocks. In ultraoligotrophic lakes.

Forming mats or hidden among other bryophytes, over peaty soils on damp ledges and crevices of rocks, sometimes along streams, or aquatic. Partial shade to full exposure.

Forming lawns or intermixed with other bryophytes in medium to rich montane fens;

grows submerged to emergent in pools or on saturated ground; usually in full sunlight.

Fen pools may dry up in late summer.

Vasculars

Elatine brachysperma ELBR5 short-seeded waterwort Growing on mudflats, shallow pools, and edges of ponds. Freely rooting in mud.

Heliotropium curassavicum HECU3 salt heliotrope Wet ground, saline flats, alkaline soils. Dried edges and centers of vernal pools. Low to moderate elevations.

Pilularia americana

PIAM

American pilwort

Rooted in mud around the edges of shallow vernal pools. In shallow water of ponds and temporary pools and reservoir margins. Semi-aquatic.

Pleuropogon oregonus

PLOR3 Oregon semaphore grass

Open, wet meadows, marshes, and riparian areas. Grows in areas of standing or flowing water early in season. Documented sites not near forested habitats. Sluggish water in depressions and sloughs, irrigation ditches.

Potamogeton diversifolius PODI diverse-leaf pondweed Aquatic; shallow creeks and ponds. Shallow water, ditches, ponds, lakes.

Potamogeton robbinsii PORO Robbins’ pondweed Shallow to deep water of ponds, lakes and slow-flowing rivers

Rotala ramosior RORA lowland toothcup Damp areas in fine silt and sand, wet swampy places, mudflats, lake and pond margins, and along free-flowing river reaches.

Utricularia minor UTMI lesser bladderwort Aquatic plant. Shallow ponds, slow-moving streams, and wet sedge or rush meadows.

Often w/bulrush and other bladderworts. Occurs in low-nutrient lakes and peat bogs.

Peatlands (Including Fens) name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Peaty soil, at high elevations. Associated with other bryophytes in tight mats. On wet soil, rocky ledges, or on wet rocks in sub alpine or alpine zones. Usually on north-facing exposures.

Peaty soil or decomposed organic detritus on damp ledges of rock outcrops and cliffs at higher elevations. Acid soil or soil-covered rocks. Not necessarily on mineral soil.

Sites receive abundant snowfall.

Calliergon richardsonii CARI70 Richardson’s calliergon moss Wet, more or less mineral- and nutrient-rich fens, ditches, shores, floating or submerged in lakes; low to high.

Campylium stellatum CAST51 Star campylium moss Intermediately mineral-rich fens, lake and river shores; low to high elevations. Often in bogs, sometimes in marshes, springs, swamps, and other moist conditions.

Bogs and fens; mostly on humus, soil covered rocks, and decaying wood in forests.

Site on Umatilla NF is in a sub-alpine meadow, surrounded by subalpine fir and spruce forest.

Polytrichum strictum

POST70 Hummock haircap

On hummocks in sphagnum bogs, wet heaths and tundra, muskeg, sedge meadows, moist alpine tundra, also on rotten stumps in wet spruce forests; low to high elevations.

narrow mushroom-headed liverwort

On soil w/little organic matter, typically on ledges or cliffs. On MAL NF in peaty fen.

Shaded creeks, seeps, splash zones. Terrestrial, on damp mineral soil; reportedly a strict calciphile, but not in Columbia Gorge. In thin soil over calcareous rocks.

Forming lawns or intermixed with other bryophytes in medium to rich montane fens;

grows submerged to emergent in pools or on saturated ground; usually in full sunlight.

Fen pools may dry up in late summer.

Splachnum sphaericum PSPS70 Dung moss Dung of large herbivores on muskeg or bog habitats at low to high elevations.

Vasculars soils. Often growing in a bed of mosses. This species tends to grow in wetter sites than the other Botrychiums.

Carex diandra

CADI4 lesser panicled sedge

Bogs and fens, floating peat mats, lake and pond shores, springs, seeps. Sometimes in brackish water. Groundwater dependent ecosystems, usually on peat soils. Wettest nonaquatic microsites in fens, pH usually between 6 and 8.

Carex lasiocarpa var. americana CALA11 slender woolly sedge Fens, bogs, lakeshores, streambanks, seepage areas, and wet meadows. In deep, organic acid soils with low to moderate nutrients. Sometimes forming floating mats.

Carex media CAME9 Scandinavian sedge

Montane to subalpine meadows, streambanks, seepage areas, bog margins. Mossy, moist forests.

Carex saxatilis CASA10 limestone or russet sedge

Fens, bogs, lakeshores, wet tundra, roadside ditches, ponds, and slow moving streams; often in shallow water. Always at high elevations. In areas with peaty soils.

Carex scirpoidea ssp. stenochlaena CASCS9 Alaskan single-spiked sedge Moist meadows, streambanks, and rocky slopes. Beside waterfalls and on seepy cliffs.

Somewhat acidic substrates.

Epilobium palustre EPPA marsh willowherb Wet meadows, shorelines, peatlands, streambanks, marshes, seeps.

Swertia perennis SWPE felwort Subalpine wet areas, stream sides, springs, wet meadows, wet alpine slopes Bogs.

Not clear if in peatlands. Moderate to high elevation in the mountains.

Warm Riparian Forests (Aspen, Cottonwood), Shrublands and Wetlands name Code Common name Habitat details

Fungi, Lichens, Liverworts, Mosses

Enthosthodon banded cord-moss, Hasselquist’s hyssop

On seasonally wet, exposed soil in seeps, or along intermittent streams. Grasslands, grassy balds, and rock outcrops. Usually hidden among grasses, other mosses, and litter.

Schistidium cinclidodonteum SCCI5 schistidium moss Grows in large clumps on wet or dry rocks, or in soil in crevices of rocks and boulders, often along intermittent streams.

Forming small turfs or cushions on soil, calcareous soil, silt, tree roots, sheltered ledges and crevices of outcrops, cliffs. Reportedly calciphile, but on acid rocks as well.

In cold forest and Populus riparian forests. Soil or rock.

Vasculars

Callitriche trochlearis

CATR2 Wheel fruited water-starwort

Wetlands. Only found at one site in the Blue Mountains, on the Malheur NF: muddy floodplain in pine forest. With Pinus contorta, Juncus balticus, Carex utriculata, C.

nebrascensis.

Calochortus longebarbatus var. peckii

CALOP4 Peck's long-bearded sego-lily

Grassy margins of wet meadows, and under pines. Wet to moist meadows. Along stream edges. Often partially shaded by Ponderosa pine. On basalt derived and volcanic ash soils.

Carex cordillerana

CACO81

cordilleran sedge

Riparian terraces, mesic forests, with grand fir and Douglas fir. Aspen forests. Rocky slopes, in leaf litter and duff. In rocky areas with juniper and mountain mahogany on Malheur NF. Near ephemeral streams in steep ravines in Hells Canyon NRA.

Carex idahoa

CAID Idaho or Parry's sedge

Flat to gently sloping meadows around headwater streams, springs. Often with Carex praegracilis and/or Poa pratensis. In ecotone between wetlands and uplands.

Subirrigated soils that dry out in summer.

Carex retrorsa CARE4 retrorse sedge Floodplains in forests, streamsides, lakeshores, swamps, wet thickets, wet meadows.

Sandy riverbanks. Sites are wet in spring, but may dry out by late summer.

Cypripedium fasciculatum

CYFA clustered lady's-slipper

Grand fir to ponderosa pine, and warm riparian forests; w/sub-alpine fir in CO.

Generally found in 60-100% shade. Often w/mtn. lady's slipper. In damp to dry sites.

Seeps/springs, riparian zones.

Diphasiastrum complanatum LYCO3 ground-cedar Dry open coniferous or mixed dry to wet forest slopes, or in riparian areas. Often on rotting logs, or in thick duff. Reportedly also in meadows and on open ridge tops.

Dracocephalum parviflorum

DRPA2 American dragonhead

Open, often moist places from the foothills to moderate elevations in the mountains;

seeds require fire or other disturbance for germination, germinating even after stand-replacing fires.

Vernally wet swales, along intermittent streams, wet depressions in moist meadows and lithosols. In slight depressions that hold snow later in the season than surrounding areas. Surrounding forest is usually ponderosa pine. Habitat and plants are dry by mid-summer.

Erythranthe inflatula

ERIN14 Disappearing monkeyflower

Vernally moist sites and fluctuating banks of intermittentstreams or pools in sagebrush-juniper zone. Amid heavy gravel and boulders at one site. In ecotone between uplands and graminoids at water's edge. Low, wet fields also.

Juncus hemiendytus var.

abjectus

JUHEA

least rush Wet banks, seeps, wet shores, wet meadows, margins of vernal pools. Sites on

Malheur are sagebrush and dry pine surrounding habitat.

Sandy soils on riverbanks, moist meadows, pond shores and edges of reservoirs.

Open, rocky and apparently dry slopes also. Open, more or Veratrum less disturbed, sandy slopes, and seeps.

Phacelia minutissima PHMI7 least or dwarf phacelia Moist, open places, streambanks, meadows, ephemerally moist swales. Vernally moist openings in ponderosa pine or Doug fir forest. Often with. In aspen in WA.

Pleuropogon oregonus

PLOR3 Oregon semaphore grass

Open, wet meadows, marshes, and riparian areas. Grows in areas of standing or flowing water early in season. Documented sites not near forested habitats. Sluggish water in depressions and sloughs, irrigation ditches.

Rorippa columbiae

ROCO3 Columbian yellowcress

Riverbanks, lake margins, wet meadows and ditches; all known sites are inundated for at least part of the year. Soil types include clay, sand, gravel, sandy silt, cobblestones and rocks.

Rotala ramosior RORA lowland toothcup Damp areas in fine silt and sand, wet swampy places, mudflats, lake and pond margins, and along free-flowing river reaches.

Under and around western juniper, ponderosa pine, and Doug fir trees; in canyons, seasonal creek drainages, and seeps/springs. Also found in vernally moist areas in ponderosa pine forests and in sage. Restricted to serpentine and ultramafics on Malheur.

Trifolium douglasii TRDO Douglas' clover Moist or mesic meadows, prairie remnants, along riparian areas along streams. In swales, along intermittent streams, and in vernally wet areas.

Warm Riparian Forests (Aspen, Cottonwood), Shrublands and Wetlands name Code Common name Habitat details

Triglochin palustris TRPA28 Slender bog arrow-grass Coastal bogs to inland meadows, mud flats, and gravelly stream margins, often where brackish or alkaline.